粉叶小檗(小檗科)的形态变异式样及分类修订

Morphological Variation Patterns of Berberis pruinosa (Berberidaceae) and Its Taxonomic Revision

摘要:

研究了云南省昆明市西山和丽江市玉龙雪山的粉叶小檗（Berberis pruinosa Franch.）叶、花瓣和果实特征的变异式样。粉叶小檗幼叶背面具明显的白色粉霜，而成熟叶，尤其是秋冬季的老叶，叶片背面的白色粉霜几乎完全脱落，并呈灰绿色。虽然粉叶小檗花瓣先端存在缺裂和全缘两种变异式样，但是这些变异式样在居群内是连续的。粉叶小檗成熟果实紫黑色，表面具显著而稳定的白色粉霜。结合文献分析和植物标本观察，将B. pruinosa var. brevifolia Ahrendt、B. pruinosa var. longifolia Ahrendt、B. pruinosa var. serratifolia Ahrendt和易门小檗处理为粉叶小檗的异名。此外，还为粉叶小檗指定了后选模式，即Delavay 493（P00716556）。

Abstract:

The variation patterns of the leaf, petal and fruit of Berberis pruinosa Franch. were studied from populations in Xishan Mountain of Kunming City and Yulongxueshan Mountain of Lijiang City, Yunnan Province, China. The abaxial surface of young leaves is densely pruinose, but the mature leaves, especially those in autumn and winter are almost abaxially glabrous and grey green, because the pruinose layers had fallen away. Although two basic variation patterns of the apex of petals are found in B. pruinosa, including entire and incised petals, however, they are consistent within populations. The ripe fruits of B. pruinosa are purplish black with a conspicuous and stable pruinose layer. In combination with studies of literatures and plant specimens, B. pruinosa var. barresiana Ahrendt, B. pruinosa var. brevifolia Ahrendt, B. pruinosa var. longifolia Ahrendt and B. pruinosa var. serratifolia Ahrendt are all treated as synonyms of B. pruinosa. In addition, a lectotype, i.e. Delavay 493 (P00716556), is designated here for B. pruinosa.