砂仁种子的解剖学和组织化学研究
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ANATOMY AND HISTOCHEMISTRY OF THE SEEDS OF AMOMUM VILLOSUM
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    摘要:

    砂仁种子包括假种皮、种皮、外胚乳、内胚乳与胚.假种皮由内表皮、外表皮及其间的6—9层薄壁细胞组成.种皮分为外种皮、中种皮与内种皮.外种皮由1层表皮细胞构成,其壁增厚并略木质化.中种皮包括各含1层细胞的下皮层和半透明细胞层与含3—5层细胞的色素层,下皮层与色素层细胞均含有红棕色素,后者的壁呈网状增厚.内种皮由1层内切向壁与径向壁非常增厚的石细胞构成.种皮表面具有许多疣状突起.它们是体积较小的表皮细胞和其内方1—3个局部膨大的下皮细胞向外突起形成的.合点区内种皮出现缺口.缺口附近石细胞增至2—3层;缺口间的合点区色素细胞整体轮廓呈喇叭状.珠孔区分化出珠孔领、孔盖与珠孔区薄壁细胞;孔盖由3—5层石细胞与2—4层色素细胞构成.内胚乳上半部具多层细胞,下半部为单层细胞.脂类物质(含挥发油)主要存在于胚和假种皮细胞中.其次为内胚乳最外层细胞与外种皮.

    Abstract:

    The seed of Amomum villosum Lour. comprises aril, seed coat, perisperm, endosperm and embryo. The aril contains outer epidermis, inner epidermis and 6-9 layers of cells between them. The seed coat can be divided into exotesta, mesotesta and endotesta. The exotesta consists of one layer of epidermal cells with thickened and slightly lignified cell walls. The mesotesta includes one layer of hypodermaal cells, one layer of translucent cells and 3-5 layers of pigment cells. The hypodermal cells and pigment cells with reticulate wall thickening are rich in plament. The endotesta is composed of one layer of sclereids and their anticlinal and inner periclinal walls are strongly thickened. The verruciform protuberances are scattered on seed coat surface. Each protuberance is constructed by 1- 3 hypodermal cells with projections outward at one end or median part of cells and their outer epidermal cells with smaller volume. A gap appears in the endotesta of the chalazal region. The number of layer of sclerids near the gap often increases to 2-3 layers. Chalazal pigment cells stuff the gap and as a mass with trumpet-shaped project inward. Micropylar collar, operculum and parenchyma cells are differentiated at micropylar region. The operculum consists of 3- 5 layers of sclereids and 2- 4 layers of pigment cells. The cndosperm has two parts, the upper part is of multilayered structure, but the lower part is with monolayer of cells. Lipids (including volatile oil) mainly present in embryo cells and several layers of cells of aril adjacent to seed coat, then in cells of the outer most layer of the endosperm and exotesta cells.

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吴七根,廖景平.砂仁种子的解剖学和组织化学研究[J].热带亚热带植物学报,1995,3(2):52~59

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